∞ generated and posted on 2016.08.29 ∞
Organisms which do not modify environments in such a way that, as a result, their own offspring would be put at a disadvantage relative to competitors.
In Ohio, the maple rees readily grow in the shade of other maple trees and thus can persist in the same general location over multiple generations and indeed indefinitely barring climate change or other catastrophes.
Note that climax species as well as ecological succession in general typically is considered primarily in terms both plant species and terrestrial environments, that is, rather than from the vantage of animals or, necessarily, aquatic environments.