Extinct tree-sized lycophytes from later in the to earlier in the period.

The surfaces of the sigillarian and likely were rather than serving as protective and probably were responsible for stiffening the trunks of these trees, that is, rather than this stiffness coming from . Sigillarians also were heterosporous.

The lepidodendrons were -related plants, like the Lepidodendron, and also contributed to the formation of , though less so than the lepidodendrons.